主要動詞

beater

Despite its high density of leopards, Malawi could previously only lay claim to a few itinerant lions in southerly Liwonde National Park. The solitary male, known as Titus, was as elusive as the literary character after which he was named; spotted in 2007 he went to ground until 2010, only to be seen once more before disappearing in 2011. "Not much of a reason for big cat buffs or travellers keen on spotting the precious Big Five (lion, rhino, buffalo, leopard, elephant) to set their compass for this southern African country.:...

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tyge318

這不是倒裝句，是不完整句，所以你當然找不到動詞。
這也是從BBC上看到的文章，是第一段的末句；為了方便說明，我把第一段完整的貼上來：

Despite its high density of leopards, Malawi could previously only lay claim to a few itinerant lions in southerly Liwonde National Park. The solitary male, known as Titus, was as elusive as the literary character after which he was named; spotted in 2007 he went to ground until 2010, only to be seen once more before disappearing in 2011. Not much of a reason for big cat buffs or travellers keen on spotting the precious Big Five (lion, rhino, buffalo, leopard, elephant) to set their compass for this southern African country....

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beater

了解
豈止滿意
簡直可以跪下來叫你一聲爺了!!!!